The lower shaft of a feather, specifically the region lacking barbs.
A pen made from a feather.
Any pen.
He picked up his quill and wrote a poem.
A sharply pointed, barbed, and easily detached needle-like structure that grows on the skin of a porcupine or hedgehog as a defense against predators.
A thin piece of bark, especially of cinnamon or cinchona, curled up into a tube.
The pen of a squid.
The plectrum with which musicians strike the strings of certain instruments.
The tube of a musical instrument.
Something having the form of a quill, such as the fold or plain of a ruff, or a spindle, or spool, upon which the thread for the woof is wound in a shuttle.